Unveiling Giganotosaurus: The Prehistoric Rival of Tyrannosaurus Rex

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Image Credit: Giganotosaurus skeleton. Fernbank Museum of Natural History in Atlanta, Georgia.

Boasting a skull as long as a man and a body the length of a bus, Giganotosaurus is among the largest predatory theropods ever discovered. Before Tyrannosaurus Rex reigned as the King of the Dinosaurs, a larger theropods dominated the prehistoric landscape. His name Giganotosaurus Carolinii.

Known as the “Giant Southern Lizard”, Giganotosaurus was a formidable predator that dominated the Southern Hemisphere. This massive theropod, a member of the Carcharodontosauridae family, hunted titanosaurs and other herbivores, establishing itself as one of the largest carnivorous dinosaurs, surpassing the mighty T-rex by 2.2 tons.

Image Credit: Comparison of the largest theropods that walked the Earth. Giganotosaurus (Green), Tyrannosaurus Rex (Blue), and Spinosaurus (Red).

The tale of Giganotosaurus began in 1993 with the discovery of a tibia jutting from the earth in Patagonia. In 1994, paleontologists revealed the unearthing of a massive new theropod. The fossilized remains comprised a partial skull, a large portion of the vertebral column, elements of the pelvis, and fragments of limb bones.

Image Credit: Giganotosaurus holotype (first Giganotosaurus skeleton found) laid in the dirt. Natural Science Museum at El Chocón, in the northwestern Argentine Patagonia.

The discovery of Giganotosaurus is important because it deepened our understanding of the Carcharodontosaurid clade. Producing some of the largest theropods to ever live such as the newly discovered Meraxes Gigas, Acrocanthrosaurus, Carcharodontosaurus, and Giganotosaurus. This clade is of further interest to dinosaur enthusiasts because it allows us to explore the upper limit of theropod size.

Image Credit: Noelle K. Moser. Alamosaurus (Titanosaur) and Tyrannosaurus Rex. Giganotosaurus preyed upon Titanosaurs during the Cretaceous. Perot Natural History Museum. Dallas, Texas.

Nature maintains a delicate balance between predators and prey. Large herbivores require equally formidable carnivores to sustain this balance. Giganotosaurus, a giant theropod, played a crucial role in the ecosystem where it lived. The real question is not whether Giganotosaurus hunted these massive herbivores, but how it did so. This article will explore the origins of the Giganotosaurus, its hunting strategies, and ultimately why it faced extinction.

Origins of Giganotosaurus:

Image Credit: Noelle K. Moser. Giganotosaurus and I. Knoxville Zoo. Knoxville, TN.

During the Mesozoic, an evolutionary arms race between herbivores and carnivores ensued. As herbivores grew larger to gain a competitive advantage, the theropods also increased in size. The Jurassic period, marking the middle era of the age of dinosaurs, witnessed a remarkable diversification in dinosaur body plans. Herbivores grew larger, and thundering across the landscape were the sauropods, the giants of the Mesozoic era, including species such as Diplodocus and titanosaurs.

Giganotosaurus belongs to the Carcharodontosauridae family, a group of theropod dinosaurs known for producing some of the largest carnivores ever to walk the earth. Besides their massive size, a distinctive characteristic of this group is streamlined narrow skulls with shark-like teeth.

Image Credit: Carchardontosaurus skull. The Carcharodontosauridae family possessed shark-like serrated teeth perfect for tearing the flesh off bones.

Teeth reveal much about a creature. By examining dinosaur teeth, we can determine their diet, hunting methods, and how they consumed their prey.

During the Jurassic, the middle period of the Mesozoic Era, there was a significant increase in size among species as a result of an evolutionary arms race between predators and prey. As herbivores grew larger, carnivores also evolved to match their size.

The Jurassic saw some of the largest and most famous herbivores – the sauropods. Species such as Diplodocus, Brachiosaurus, Supersaurus, and Camarasaurus.

Image Credit: Comparison Graphic of some of the longest Sauropods that lived during the Jurassic.

Counterparts to these lumbering giants, were the carnivores of the Jurassic, relatives of Giganotosaurus such as Tyrannotitan, Lusovenator, Siamraptor, and Acrocanthrosaurus.

Image Credit: An overview of the paleofauna that inhabited the Southern Hemisphere during the Mesozoic era includes theropods such as Mapusaurus (red) and Meraxes gigas (dark blue), which are carcharodontosaurs and related to Giganotosaurus.

Inhabiting the Southern Hemisphere, the relatives of Giganotosaurus, known as primitive Carcharodontosaurs, evolved into increasingly larger theropods in response to the growing size of the herbivores they preyed upon. By the end of the Jurassic and into the Early Cretaceous, the Carcharodontosauridae family comprised some of the largest carnivorous dinosaurs to have ever walked the Earth.

Image Credit: Noelle K. Moser. Life-size reconstruction of Giganotosaurus. Knoxville Zoo. Knoxville, Tennessee.

Giganotosaurus represented the culmination of an evolutionary arms race, standing as the pinnacle of the Carcharodontosauria clade.

How Giganotosaurus Hunted and Killed Prey:

Analysis of the leg bones of Giganotosaurus shows that this theropod was not built for speed, but it didn’t need to be. Although it was slower than the swift herbivores, Giganotosaurus preyed on the more ponderous sauropods, known as titanosaurs.

The titanosaurs were the last surviving group of long-necked sauropods, thriving at the time of the Chicxulub Impact at the end of the Cretaceous that ended the age of the dinosaurs. This group includes some of the largest land animals known to have ever existed, such as Argentinosaurus.

Image Credit: Argentinasaurus and Giganotosaurus displayed together. Fernbank Museum of Natural History. Atlanta, Georgia.

Titanosaurs lived by one rule, get big and get big fast. From the moment of hatching, sauropods like Argentinasaurus were eating machines. Dining on leaves and hard fibrous vegetation, a herd of titanosaurs could defoliate an area in a few days.

Large guts and hard-to-digest food allowed for a slow release of energy over time. This superpower aided in the ability of these sauropods to reach full size in less than ten years. Once fully grown, an adult Argentinasaurus was 128 ft long, 65 ft tall, and weighed 65 to 82 tons. This sheer size alone was enough to detour many theropods from making a meal out of these massive herbivores. Traveling in herds combined with size officially removed them from the menu.

Hunting a herd of titanosaurs was perilous. A single misstep can result in one of these colossal herbivores crushing an overzealous theropod, leading to instant death. Considering this risk, the question is not whether Giganotosaurus hunted titanosaurs, but rather how they accomplished such a feat.

Much like the enigmas posed by extinct species, the most effective way to address these questions is by examining the present. Observing lions as they hunt a herd of wildebeests, we see the predators collaborate to disperse the group, targeting the smaller, ill, or weakest members for an easier kill. A lion understands that to attack the largest, strongest, or healthiest would be, at best, a perilous endeavor. This logic can be similarly applied to Giganotosaurus.

Traveling herds exhibit remarkable organization. The young and subadults are positioned centrally, while the robust and healthy adults encircle them, forming a protective barrier. Typically, the elderly or injured members trail behind, comprising the rear guard as the herd moves across the terrain.

Understanding herd dynamics, a hunting Giganotosaurus would likely approach the herd from behind, targeting the weaker Argentinasaurus individuals. Despite not being in their prime, these titanosaurs remained formidable, capable of inflicting fatal injuries. It is probable that for these reasons, Giganotosauruses would hunt in packs, coordinating their efforts to take down one of these colossal creatures.

Evidence from the teeth of Giganotosaurus suggests that, unlike the bone-crushing bite of Tyrannosaurus Rex, Giganotosaurus had teeth better suited for slicing off flesh from its prey. Packs of Giganotosaurus would alternate in biting and slashing their prey, aiming to keep it moving and bleeding. The hunting strategy was to exhaust the prey through blood loss, fatigue, and infections caused by the theropods attacks, leading to the titanosaur’s eventual collapse under its own weight.

Trace fossils provide definitive evidence of theropod hunting strategies located along the Paluxy River near Glen Rose, Texas, USA. Here, a dramatic narrative of a dinosaur hunt is etched into the stone.

Image Credit: Peluxe River in Glenn Rose, Texas. Footprints from a Cretaceous hunt involving Acrocanthrosaurus and Saurposeidon recorded in stone.

120 million years ago, on a muddy Cretaceous floodplain, the dynamics of dinosaur relationships were immortalized in stone. A herd of colossal sauropods lumbered along a waterway, stalked closely by a large carnivore. The pursuing theropod was focused, intent on the hunt.

Following behind the herd, slightly to the left, the theropod’s tracks indicate that the hunter rhythmically trailed the lumbering sauropods. Then the theropod’s footprints show that the hunter suddenly skipped a few steps, meaning only one thing, an attack.

Most of the trackway was removed. It is now preserved and displayed at The American Museum of Natural History in New York. Some of the trackway still remains submerged under the Paluxy River near Glen Rose, Texas.

Image Credit: Trackway from the Peluxy River on display at The American Museum of Natural History in New York.

Giganotosaurus Extinction:

Giganotosaurus lived during the Late Cretaceous period, specifically in the Cenomanian stage, approximately 99.6 to 97 million years ago. The reasons for its extinction are not definitive, but fossil records suggest several plausible scenarios. During the latter part of the Cretaceous, environmental changes due to plate tectonics posed survival challenges for Giganotosaurus and other Carcharodontosaurids.

Additionally, around 30 million years ago, Tyrannosaurs emerged as the dominant carnivores, with Abelisaurs prevailing in the Southern Hemisphere and Tyrannosaurus Rex in the northern. It is conceivable that Giganotosaurus was outcompeted by these more adaptable theropods, leading to a gradual decline and eventual extinction.

After the extinction of the last of the Carcharodontosaurs, Giganotosaurus lost its dominance, allowing the Tyrannosaurus and the formidable Tyrannosaurus Rex to rise as the King of the Dinosaurs until 66 million years ago when the age of the dinosaurs came to an end.

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Image Credit: Noelle K. Moser. Me peering through the fenestra of Tyrannosaurus Rex MOR 555. Cincinnati Natural History Museum. Cincinnati, Ohio.

Resources:

Johnson-Ransom, Evan. Dinosaur World: Over 1,200 Amazing Dinosaurs, Famous Fossils, and the Latest Discoveries from the Prehistoric Era. Applesauce Press. Kennebunkport, Maine. 2023.

Keiron, Pim. Dinosaurs The Grand Tour: Everything Worth Knowing About Dinosaurs from Aardonysx to Zuniceratops. The Experiment. New York, NY. 2019.

My visit to Natural History Museums across the nation.

Allosaurus: Butcher of the Jurassic

Image Credit: Noelle K. Moser. Allosaurus: Houston Museum of Natural History

One-hundred and fifty million years ago, long before T-Rex reigned as the tyrant king of the dinosaurs, another fearsome theropod ruled the land his name, Allosaurus (AL-oh-SORE-us) Fragilis. Spanning a dynasty from 150 million to 144 million years ago during the late Jurassic period, Allosaurus is famous from the Cleveland-Lloyd Quarry in Utah, part of the Morrison Formation and one of the best-known ancient dinosaur graveyards in the world where the remains of 44 Allosaurus specimens lay entombed. Victims of a predator trap, the Allosaurus specimens from this Quarry are some of the actors in an epic prehistoric tragedy that unfolded 150 million years ago; this is the tale of that fateful day.

Image Credit: Allosaurus skeleton from the Cleveland-Lloyd Quarry.

Cleveland-Lloyd Quarry:

Before we can understand this epic tale and the part that Allosuaurs plays, we must look at the stage where this drama unfolds. The Cleveland-Lloyd Dinosaur Quarry well known for containing the densest concentration of Jurassic dinosaur fossils ever found, is a paleontological site located near Cleveland, Utah in the San Rafael Swell, a part of the geological layers known as the Morrison Formation.

Morrison Formation:

Image Credit: Morrison Formation in Utah

The Morrison Formation is a distinctive sequence of Upper Jurassic sedimentary rock found in the western United States, which has been the most fertile source of dinosaur fossils in North America and covers an area of 600,000 square miles. Composed of mudstone, sandstone, siltstone, and limestone with a light gray, greenish, and red sediment, the Morrison Formation was a Jurassic waterway shortly after Lauraisa split into the continents of North America and Eurasia.

Image Credit: Map of the continents 150 million years ago during the Jurassic Period.

As North America moved north and passed through the subtropical regions, the deposits from drainage basins carried by streams and rivers became the Morrison Formation. Above is a map of the Earth as the contents traversed the globe due to plate tectonics.

Image Credit: Cleveland-Lloyd Quarry map of Jurassic Dinosaurs bones.

Above is a map of all the dinosaur bones found preserved within the Cleveland-Lloyd Quarry. Looking at this grapic two things are abundantly clear.

First, the amount of carnivores present at the Cleveland-Lloyd Quarry is staggering. Indicated in green, the theropods present are predominately Allosaurus. Accompanying Allosaurus are the remains of Ceratosaurus (SERR-at-oh-SORE-us) – the most dragon like-dinosaur known and relative of Abelisaurus, Stokesaurus – the first tyrannosauroids and primitive ancestor of Tyrannosaurus Rex, Mashosaurus (MARSH-o-SORE-us), Torvosaurus (TORE-voh-SORE-us) the biggest carnivore of its time – as big as Allosaurs and heavier too, and Tanycolagreus – a primitive coelurosaurian theropod.

Second, the herbivores present are fewer but noteworthy. In yellow are the remains of several massive sauropods, Camarasaurus (CAM-a-ra-SORE-us) – 25.4 tons and Brontosaurus Excelsus (BRONT-oh-SORE-us) – 18 tons. Denoted in blue are the remains of an Ornithopod Camptosaurus (CAMP-toe-SORE-us), and indicated in red are the remains of several Thyreophora or armored dinosaurs known as Stegosaurus.

Image Credit: A Barosaurus defending itself against a pair of Allosaurus Thereopods.

Upon looking at the remains encompassing a wide variety of different dinosaurs, it is clear that something unusual attracted the assemblage of paleofauna to this location. Carnivores outnumbering herbivores points to scenes in which predators flocked to the site. What drew the diverse number of theropods to this site 150 million years ago? This Jurassic crime scene preserved in stone – the Cleveland-LLoyd Quarry – is a predator trap.

What is a Predator Trap? How Does it work?

predator trap results from a natural hazard where prey animals become trapped or incapacitated, and the predators attracted to the struggling prey suffer the same fate. As more and more animals become ensnared in the trap, additional predators and scavenges become attracted to this mounting accumulation of carrion until a wide variety of animals are tragically caught and killed by the natural hazard. The predator trap can appear many times, accumulating countless victims over millions of years.

Unlike the dry landscape today, 150 million years ago, the Cleveland-Lloyd Quarry was a freshwater lake, a source of water that drew many dinosaurs to its shores. Much like Africa today, this area had wet and dry seasons. The wet seasons, with torrential rains and floods, would fill the landscape with water. As the dry season approached, the water would dry up; dense mud would then take the place of the water. It is this weather cycle that sets the stage for what happens next.

How did the Cleveland-Lloyd Quarry Drama Unfold?

While it’s impossible to say exactly how the events unfolded at the Cleveland-Lloyd Quarry 150 million year ago, fossil evidence suggests a probable interpretation of that fateful day. Brad Cypy. (2014, July 13) Jurassic Fight Club: Bloodies Battle {Video File}. YouTube

Cleveland-Lloyd Quarry. 9:00 am – Act 1; Scene 1. Adult Stegosaurus and Offspring Enters, Stage Right.

Image Credit: Stegosaurus

A Stegosaurus and her young searching for water approach the shrinking lake bed. The hardened upper crust buckles under their massive weight, causing their feet to sink into the soft muck below and become affixed. The cries of distress from the Stegosaurus pair attract a lone carnivore, Ceratosaurus (SERR-at-oh-SORE-us).

Cleveland-Lloyd Quarry. 9:30 am. – Act 1; Scene 2. Ceratosaurus Enters Stage Left.

Image Credit: Ceratosaurus

The fossil record shows that a lone Ceratosaurus was the first to arrive at the scene at Cleveland-Lloyd Quarry. One of the rarest killers, Ceratosaurus, is 13 feet tall, 20 feet from snout to tail, and weighs 3,000 pounds, a mid-size predator in the Late Jurassic. Ceratosaurus may not have been massive, but his oversized teeth that worked like a meat slicer made him a fearsome beast. (Pim, 2019)

Ceratosaurus underneath a Stegosaurus. National Museum of Natural History. Washington, D.C.

Ceratosaurus has the most wicked teeth of any predator that walked the Earth because he has two different kinds of teeth, a trait rare in theropods. The upper teeth were long and blade-like, and the lower teeth were conical and more powerful. Ceratosaurs would use the lower jaws to grab ahold of flesh and bring down the top teeth like a meat slicer cutting a chunk out of his prey. Ceratosaurus’s bite allowed him to compensate for two weaknesses: diminished forearm claws and smaller leg muscles, reducing speed and agility. Not equipped for battle, Ceratosaurus is made to slash and dash. Dinosaurs: The Complete, Up-To-Date Encyclopedia. (Holtz & Rey. 2007).

Observing the Stegosaur pair trapped in the mire, Ceratosaurus attacks the smaller Stegosaurus. Taking massive chunks out of the struggling Stegosaurus pair.

Cleveland-Lloyd Quarry. 10:00 am. – Act 1; Scene 3. A small group of Allosaurus Approach the Lakebed. Center Stage.

The most common dinosaur found at the Cleveland-Lloyd Quarry site is Allsoaurus. Allosaurus predates Tyrannosaurus Rex by 80 million years and is the first giant theropod to roam North America.

Image Credit: Allosaurus Attacking Stegosaurus Pair.

The name Allosaurus means “Other Reptile”. At its time, there was nothing as fearsome as Allosaurus. The first of the giant meat-eating dinosaurs, this carnivore was Tyrannosaurus Rex before there was Tyrannosaurus Rex. Possessing a rare trait in theropods, Allosaurus had “Freddie Krueger” like hands with three fingers armed with a long, sharp sickle claw used to slash and tear flesh. 13 feet tall, 40 feet from snout to tail, and weighing up to 8,000 pounds (4 tons), Allosaurus is the butcher of the Jurassic. (Pim, 2019).

Allosaurus, equipped to take on big game hunted sauropods, some of the largest animals the planet has ever seen during the Late Jurassic. A long, rigid tail provided excellent balance. Coupled with massive leg muscles, Allosaurus is a powerful, fast runner with incredible agility. Dinosaurs: The Most Complete Up-To-Date Encyclopedia (Holtz & Rey, 2007).

Image Credit: Endocast of Allosaurus Brain

Endocasts of the Allosaurus brain reveal that in addition to being the first giant theropod, Allosaurus was also the smartest carnivore in the Late Jurassic. Allosaurus was capable of advanced social behaviors such as cooperative hunting and planning. Like modern-day birds, Allosaurus was very social and lived in small groups.

A roaming group of Allosaurus arrive at the dried lake bed, attracted by the sound of disruption and the smell of fresh blood. The Allosaurus see the Stegosaurus pair stuck in the thick mire and a single Ceratosaurus. The fossil record shows that a single Ceratosaurus was present at the Cleveland-Lloyd Quarry, its bones crushed by another dinosaur.

Cleveland-Lloyd Quarry. 10:30 am – Act 1; Scene 4. The Allosaurus Kill the Ceratosaurus and feed on the Stegosaurus Pair.

Attempting to defend its meal against the Allosaurus group, the Ceratosaurus has an altercation with the Allosaurus. Small and not robust like the much more derived Allosaurus, the Ceratosaurus is out-competed, crushed, and killed by better-equipped theropods.

Splade toes allow the theropods to walk on top of the hardened crust of the dried lake bed without sinking into the muck below. Circling the herbivores, the Allosaurus group continues to feed on the larger struggling Stegosaurus. Armed with plates along its back and spikes at the end of its tail, the Stegosaurus uses all its defenses to the best of its ability. The fossil records show that one of the Allosaurus has pathologies indicative of a Stegosaur strike by the spikes on its tail. The other Allosaurus, underestimating the Stegosaurus’s ability, takes a step back when another group of dinosaurs enters the scene.

Splayed toes of theropods allow them to distribute their weight over larger areas.

Cleveland-Lloyd Quarry. 11:00 am. – Act 2; Scene 1. Camarasaurus Herd Enter, Stage Left.

Image Credit: Camarasaurus Herd

Traversing the Jurassic landscape on yearly migrations, a traveling herd of Camarasaurus severely parched arrived at the water source. Due to the ongoing drought resulting from the dry season, the much-needed water had transitioned to a thick, soupy mud. Motivated by desperate thirst, the Camarasaurus herd approached the moist, thick muck.

Witnessing the unfolding bloody scene, driven by intense thirst, the leading Camasasaurus cautiously approaches the dried lake bed. Victim of his weight, the Camasaurus breaks through the hardened top layer and becomes entrapped by its weight, and the 25-ton Camarasaurus’s feet sink deep into the silt. Unable to free its feet, the Camarasaurus cried in distress. The Allosaurus, determining that attacking the Stegosaurus is too risky, sees the Camarasaurus as an easier target and approaches the struggling titan.

Unable to free its feet, the Camarasaurus cried in distress. Another group of nearby Allosaurus heard the cries of anguish and approached the scene. Upon observing the entrapped Sauropod, the carnivores assumed a free meal was at hand, hot-footed toward the struggling prey.

Cleveland-Lloyd Quarry. 11:30 am. – Act 2; Scene 2. The Noise, Smell of Blood, and Carnage Attract More Predators.

Adding to the chorus of distressing calls of anguish, the entrapped herbivores, and battling carnivores attract more predators to the scene.

The fossil record of this event shows that under the feet of the Camarasaurus lies the crushed body of an Allosaurus. At some point during the struggle, the Camarasaurus freed its front legs and reared up on its hind legs, crushing an Allosaurus running towards it.

More carnivores arrive, attracted by the carnage as the battle rages. Allosaurus, the most common animal at this site, fought and killed each other as they competed for the carcasses. As more carnivores became trapped in the thick mire, the predator trap claimed more victims. The noise at Cleveland-Lloyd Quarry on this day 150 million years ago would have been deafening. It is evident from the events recorded in stone that the land flowed with blood.

~ End Scene ~

Image Credit: Carnivores Fighting Over Carrion at Clevenland-Lloyd Quarry.

This Jurassic crime scene is a resource for dinosaur bones and allows us to understand how these animals relate to each other. The largest dinosaur graveyard, Cleveland-Lloyd Quarry, is still revealing its secrets.

I am a multi-disciplinary writer, blogger, and web content creator. Please visit my writing portfolio and other blogs if you like this post.

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Allosaurus and I. National Museum of Natural History in Washington, D.C.

Resources:

Holtz, Thomas. R. Jr. & Rey, Luis V. Dinosaurs. The Most Complete, Up-To-Date Encyclopedia. New York, Random House, 2007.

Pim, Keiron & Pastori, Fabio. Dinosaurs The Grand Tour. New York, The Experiment, 2019.

Ransom-Johnson, Evan & Csotonyi, Julius. Dinosaur World. Kennebunkport, Maine, Applesauce Press, 2023